The Laughingthrushes are the genus Garrulax of the large Old World babbler family of passerine birds. Chordates ( Phylum Chordata) are a group of Animals that includes the Vertebrates together with several closely related Invertebrates Birds ( class Aves) are bipedal endothermic ( Warm-blooded) Vertebrate animals that lay eggs. The Old World babblers or timaliids are a large family of mostly Old World Passerine Birds They are rather diverse in size and coloration René Primevère Lesson (March 20 1794 - April 28 1849 was a French surgeon and naturalist. The Old World babblers or timaliids are a large family of mostly Old World Passerine Birds They are rather diverse in size and coloration Birds ( class Aves) are bipedal endothermic ( Warm-blooded) Vertebrate animals that lay eggs. They occur in tropical Asia, with the greatest number of species occurring in the Himalaya and southern China. China ( Wade-Giles ( Mandarin) Chung¹kuo² is a cultural region, an ancient Civilization, and depending on perspective a National
These are rangy, medium-sized, floppy-tailed landbirds with soft fluffy plumage. These birds have strong legs and are quite terrestrial. This group is not strongly migratory, and most species have short rounded wings, and a weak flight. Bird migration refers to the regular seasonal journeys undertaken by many species of Birds Bird movements include those made in response to changes in food availability In Biology, a species is one of the basic units of Biological classification and a Taxonomic rank.
A few, like the Streaked Laughingthrush occur in fairly open habitats, but most are jungle species, difficult to observe in the dense vegetation they prefer. UserPolbot. -->The Streaked Laughingthrush ( Garrulax lineatus) is a species of Bird in the Timaliidae
Like other babblers, these are noisy birds, and the characteristic laughing calls are often the best indication that these birds are present. As with other babbler species, they frequently occur in groups of up to a dozen, and the rainforest species like the Ashy-headed Laughingthrush often occur in the mixed feeding flocks typical of tropical Asian jungle. The Ashy-headed Laughingthrush, Garrulax cinereifrons, is an Old World babbler.
Systematics
As with some other babblers, this genus has turned out to be highly polyphyletic. In Phylogenetics, a Taxon is polyphyletic ( Greek for "of many races" if the trait its members have in common evolved separately in different Rather than forming a natural evolutionary group of closest relatives, the laughingthrushes are spread about the entire babbler family. eVolution is the third Album by eLDee, it was due to be released in 2008 This is quite apparent by their diverse morphology and presence of several groups with distinct color patterns, but the rarity of several species and the sheer number of taxa have hitherto prevented a thorough revision. The term morphology in Biology refers to the outward appearance ( Shape, Structure, Colour, Pattern) of an Organism A taxon (plural taxa) or taxonomic unit, is a name designating an organism or a group of Organisms In Biological nomenclature according to Before this paraphyly was realized to be as dramatic as it is, it was generally assumed that the laughingthrushes were a clade comprising several subgenera. A clade is a taxonomic group comprising a single Common ancestor and all the descendants of that ancestor In Biology, a subgenus is a Taxonomic rank directly below Genus.
This has much confounded further research: most molecular phylogeny studies, for example, used only the White-crested Laughingthrush as an "example" of this "genus", because specimens were readily available. The White-crested Laughingthrush, Garrulax leucolophus is an Old World babbler. Morphological studies, on the other hand, had usually more material at their disposal, but these too were generally conducted under a false assumption of monophyly. A 2003 study[1], analyzing mtDNA cytochrome b and 12S/16S rRNA data of a number of laughingthrushes, did establish however that there is no such thing as a "representative" laughingthrush. Mitochondrial DNA ( mtDNA) is the DNA located in Organelles called mitochondria. Cytochrome b/b6 is main subunit of transmembrane cytochrome bc1 and b6f complexes Consequently most of the work regarding the evolution of this assemblage needs to be evaluated anew and if necessary even redone. Indeed, the taxon Garrulax would need to be restricted to the type species (Rufous-fronted Laughingthrush) and its closest relatives, but the species generally considered the most likely candidates have not been comprehensively studied in recent times[2]. In Taxonomy, a type species is the species that originally defined a genus. UserPolbot. -->The Rufous-fronted Laughingthrush ( Garrulax rufifrons) is a species of Bird in the
Species list
The list below uses the treatment of the laughingthrushes by Birds of South Asia[3] and the Handbook of Birds of the World[4] as a baseline, acknowledging the polyphyly of Garrulax and attempting to reorganise it into monophyletic genera. Birds of South Asia redirects here For a list of the birds of South Asia see List of birds of South Asia. HBW_-_Taxonomygif|right|frame|(400 × 258 pixels file size38KBMIME typeimage/gif|HBW-Page on Taxonomy]]HBW-accounts_8-082-083_copia A clade is a taxonomic group comprising a single Common ancestor and all the descendants of that ancestor
This list is by no means the last word on the subject though, and it is very likely that some of the new groups will be revised, expanded or split as more species are studied in the future; additional laughingthrushes are presently elevated to species status and even hitherto unknown taxa are discovered at a rate of about 1-2 per year. Some proposed small or monotypic genera on the other hand might ultimately be merged with others for convenience if they turn out to be sister taxa. Monotypic is an adjective that refers to a taxonomic group with only one type: In Botany, "monotypic" means that a Taxon has only Cladistics is the hierarchical classification of Species based on evolutionary ancestry All in all, the resolution of the laughingthrush assemblage's taxonomy depends on a resolution of the Old World babblers' systematics in general. Taxonomy is the practice and science of classification The word comes from the Greek, taxis (meaning 'order' 'arrangement' and, nomos Biological systematics is the study of the diversity of Life on the planet Earth both past and present and the relationships among living things through time
- Genus Garrulax sensu stricto - Close to Chinese Babax (Babax lanceolatus)?
- White-crested Laughingthrush, Garrulax leucolophus
- Sumatran Laughingthrush, Garrulax bicolor
- Lesser Necklaced Laughingthrush, Garrulax monileger
- Greater Necklaced Laughingthrush, Garrulax pectoralis
- White-throated Laughingthrush, Garrulax albogularis
- Rufous-crowned Laughingthrush, Garrulax ruficeps - formerly in G. The White-crested Laughingthrush, Garrulax leucolophus is an Old World babbler. The Laughingthrushes are the genus Garrulax of the large Old World babbler family of Passerine Birds They occur in tropical UserPolbot. -->The Chinese Babax ( Babax lanceolatus) is a species of Bird in the Timaliidae The White-crested Laughingthrush, Garrulax leucolophus is an Old World babbler. The Sumatran Laughingthrush, Garrulax bicolor, is an Old World babbler. UserPolbot. -->The Lesser Necklaced Laughingthrush ( Garrulax monileger) is a species of Bird in the UserPolbot. -->The Greater Necklaced Laughingthrush ( Garrulax pectoralis) is a species of Bird in the UserPolbot. -->The White-throated Laughingthrush ( Garrulax albogularis) is a species of Bird in the albogularis
- Masked Laughingthrush, Garrulax perspicillatus
- Rufous-fronted Laughingthrush, Garrulax rufifrons
- Ashy-headed Laughingthrush, Garrulax cinereifrons
- Sunda Laughingthrush or Grey-and-brown Laughingbird, Garrulax palliatus
- incertae sedis - Garrulax or Dryonastes?
- Grey Laughingthrush, Garrulax/Dryonastes maesi
- Rufous-cheeked Laughingthrush, Garrulax/Dryonastes castanotis - formerly in G. UserPolbot. -->The Rufous-fronted Laughingthrush ( Garrulax rufifrons) is a species of Bird in the The Ashy-headed Laughingthrush, Garrulax cinereifrons, is an Old World babbler. UserPolbot. --> The Sunda Laughingthrush ( Garrulax palliatus) is a species of Bird in the Timaliidae UserPolbot. -->The Grey Laughingthrush ( Garrulax maesi) is a species of Bird in the Timaliidae /D. maesi
- Black-hooded Laughingthrush, Garrulax/Dryonastes milleti
- Kon Tum Black-hooded Laughingthrush, Garrulax/Dryonastes milleti sweeti
- Cambodian Laughingthrush, Garrulax/Dryonastes ferrarius - formerly in G. UserPolbot. -->The Black-hooded Laughingthrush ( Garrulax milleti) is a species of Bird in the Timaliidae /D. strepitans
- White-necked Laughingthrush, Garrulax/Dryonastes strepitans
- Proposed genus Dryonastes
- Grey-sided Laughingthrush, Dryonastes caerulatus
- Rusty Laughingthrush, Dryonastes poecilorhynchus
- Buffy Laughingthrush, Dryonastes berthemyi - formerly in D. UserPolbot. -->The White-necked Laughingthrush ( Garrulax strepitans) is a species of Bird in the UserPolbot. -->The Grey-sided Laughingthrush ( Garrulax caerulatus) is a species of Bird in the UserPolbot. -->The Rusty Laughingthrush ( Garrulax poecilorhynchus) is a species of Bird in the poecilorhynchus
- Black-throated Laughingthrush, Dryonastes chinensis
- Chestnut-backed Laughingthrush, Dryonastes nuchalis
- Rufous-vented Laughingthrush, Dryonastes gularis
- Wynaad Laughingthrush, Dryonastes delesserti
- Blue-crowned Laughingthrush or Courtois's Laughingbird, Dryonastes courtoisi - formerly in D. UserPolbot. -->The Black-throated Laughingthrush ( Garrulax chinensis) is a species of Bird in the UserPolbot. --> The Chestnut-backed Laughingthrush ( Garrulax nuchalis) is a species of Bird in the UserPolbot. -->The Rufous-vented Laughingthrush ( Garrulax gularis) is a species of Bird in the UserPolbot. -->The Wynaad Laughingthrush ( Garrulax delesserti) is a species of Bird in the Timaliidae The Blue-crowned Laughingthrush or Courtois's Laughingbird ( Garrulax courtoisi) is an enigmatic species of Bird in the Timaliidae galbanus
- Yunnan Blue-crowned Laughingthrush, Dryonastes courtoisi simaoensis
- Yellow-throated Laughingthrush, Dryonastes galbanus
- White-cheeked Laughingthrush, Dryonastes vassali
Spectacled Laughingthrush
Garrulax mitratus/Rhinocichla mitrata
- Rufous-necked Laughingthrush, Dryonastes ruficollis
- Proposed genus Melanocichla
- Black Laughingthrush, Melanocichla lugubris
- Bare-headed Laughingthrush, Melanocichla calva - formerly in M. UserPolbot. --> The Yellow-throated Laughingthrush ( Garrulax galbanus) is a species of Bird in the UserPolbot. -->The White-cheeked Laughingthrush ( Garrulax vassali) is a species of Bird in the UserPolbot. -->The Rufous-necked Laughingthrush ( Garrulax ruficollis) is a species of Bird in the UserPolbot. -->The Black Laughingthrush ( Garrulax lugubris) is a species of Bird in the Timaliidae UserPolbot. -->The Bare-headed Laughingthrush ( Garrulax calvus) is a species of Bird in the Timaliidae lugubris
- Proposed genus Rhinocichla - Chestnut-capped Laughingthrushes
- Spectacled Laughingthrush, Rhinocichla mitrata
- Chestnut-hooded Laughingthrush, Rhinocichla treacheri - formerly in R. UserPolbot. -->The Chestnut-capped Laughingthrush ( Garrulax mitratus) is a species of Bird in the mitrata
- Proposed genus Grammatoptila
- Proposed genus Stactocichla
- Spot-breasted Laughingthrush, Stactocichla merulina
- Orange-breasted Laughingthrush, Stactocichla annamensis - formerly in S. UserPolbot. -->The Striated Laughingthrush ( Garrulax striatus) is a species of Bird in the Timaliidae The Chinese Hwamei or Melodious Laughingthrush ( Leucodioptron canorum, formerly Garrulax canorus) is a Passerine Bird of eastern UserPolbot. -->The Spot-breasted Laughingthrush ( Garrulax merulinus) is a species of Bird in the merulina
- Proposed genus Leucodioptron
- Chinese Hwamei, Leucodioptron canorum
- Taiwan Hwamei, Leucodioptron taewanum - formerly in L. The Chinese Hwamei or Melodious Laughingthrush ( Leucodioptron canorum, formerly Garrulax canorus) is a Passerine Bird of eastern The Taiwan Hwamei ( Leucodioptron taewanum, formerly Garrulax taewanus) is a Passerine Bird in the Old World babbler family Timaliidae canorum
- Proposed genus Strophocincla
- Black-chinned Laughingthrush, Rufous-breasted Laughingthrush or Nilgiri Laughingthrush, Strophocincla cachinnans
- Grey-breasted Laughingthrush, Strophocincla jerdoni
- Kerala Laughingthrush, Strophocincla (jerdoni) fairbanki
- Bhutan Laughingthrush, Strophocincla imbricata - formerly in S. UserPolbot. -->The Streaked Laughingthrush ( Garrulax lineatus) is a species of Bird in the Timaliidae The Rufous-breasted, Black-chinned or Nilgiri Laughingthrush, Trochalopteron cachinnans, is a species of Laughingthrush endemic to UserPolbot. -->The Grey-breasted Laughingthrush ( Garrulax jerdoni) is a species of Bird in the UserPolbot. -->The Streaked Laughingthrush ( Garrulax lineatus) is a species of Bird in the Timaliidae lineata
- Streaked Laughingthrush, Strophocincla lineata
- Striped Laughingthrush, Strophocincla virgata
- Proposed genus Pterorhinus
- incertae sedis - Ianthocincla or Trochalopteron?
- Proposed genus Trochalopteron
- Scaly Laughingthrush, Trochalopteron subunicolor
- Blue-winged Laughingthrush, Trochalopteron squamatum
- Brown-cheeked Laughingthrush or Prince Henry's Laughingthrush, Trochalopteron henrici
- Elliot's Laughingthrush, Trochalopteron elliotii
- Variegated Laughingthrush, Trochalopteron variegatum
- White-whiskered Laughingthrush, Trochalopteron morrisonianum
- Black-faced Laughingthrush, Trochalopteron affine
- Chestnut-crowned Laughingthrush, Trochalopteron erythrocephalum
- Assam Laughingthrush, Trochalopteron chrysopterum - formerly in T. UserPolbot. -->The Streaked Laughingthrush ( Garrulax lineatus) is a species of Bird in the Timaliidae UserPolbot. --> The Striped Laughingthrush ( Garrulax virgatus) is a species of Bird in the UserPolbot. -->The White-browed Laughingthrush ( Garrulax sannio) is a species of Bird in the Timaliidae UserPolbot. -->The Plain Laughingthrush ( Garrulax davidi) is a species of Bird in the Timaliidae The Agoutis of the genus Dasyprocta are locally known as "cutias" UserPolbot. -->The Arrow-marked Babbler ( Turdoides jardineii) is a species of Bird in the Timaliidae UserPolbot. -->The Snowy-cheeked Laughingthrush ( Garrulax sukatschewi) is a species of Bird in the UserPolbot. --> The Moustached Laughingthrush ( Garrulax cineraceus) is a species of Bird in the UserPolbot. -->The Rufous-chinned Laughingthrush ( Garrulax rufogularis) is a species of Bird in the UserPolbot. -->The Chestnut-eared Laughingthrush ( Garrulax konkakinhensis) is a species of Bird in the UserPolbot. -->The Barred Laughingthrush ( Garrulax lunulatus) is a species of Bird in the Timaliidae UserPolbot. -->The White-speckled Laughingthrush ( Garrulax bieti) is a species of Bird in the Timaliidae UserPolbot. -->The Giant Laughingthrush ( Garrulax maximus) is a species of Bird in the Timaliidae UserPolbot. -->The Spotted Laughingthrush ( Garrulax ocellatus) is a species of Bird in the Timaliidae UserPolbot. -->The Brown-capped Laughingthrush ( Garrulax austeni) is a species of Bird in the Timaliidae The Red-winged Laughingthrush ( Garrulax formosus) is a Passerine Bird belonging to the Laughingthrush genus Garrulax in the Old UserPolbot. -->The Scaly Laughingthrush ( Garrulax subunicolor) is a species of Bird in the Timaliidae UserPolbot. -->The Blue-winged Laughingthrush ( Garrulax squamatus) is a species of Bird in the UserPolbot. -->The Brown-cheeked Laughingthrush ( Garrulax henrici) is a species of Bird in the UserPolbot. -->The Elliot's Laughingthrush ( Garrulax elliotii) is a species of Bird in the Timaliidae UserPolbot. --> The Variegated Laughingthrush ( Garrulax variegatus) is a species of Bird in UserPolbot. -->The White-whiskered Laughingthrush ( Garrulax morrisonianus) is a species of Bird in the UserPolbot. -->The Black-faced Laughingthrush ( Garrulax affinis) is a species of Bird in the Timaliidae UserPolbot. -->The Chestnut-crowned Laughingthrush ( Garrulax erythrocephalus) is a species of Bird in UserPolbot. -->The Chestnut-crowned Laughingthrush ( Garrulax erythrocephalus) is a species of Bird in erythrocephalum
- Silver-eared Laughingthrush, Trochalopteron melanostigma - formerly in T. erythrocephalum
- Malayan Laughingthrush, Trochalopteron peninsulae - formerly in T. erythrocephalum
- Golden-winged Laughingthrush, Trochalopteron ngoclinhensis
- Collared Laughingthrush, Trochalopteron yersini
- Red-winged Laughingthrush, Trochalopteron formosum
- Red-tailed Laughingthrush, Trochalopteron milnei
References
- Cibois, Alice (2003a): Mitochondrial DNA Phylogeny of Babblers (Timaliidae). UserPolbot. -->The Golden-winged Laughingthrush ( Garrulax ngoclinhensis) is a species of Bird in the UserPolbot. -->The Collared Laughingthrush ( Garrulax yersini) is a species of Bird in the Timaliidae The Red-winged Laughingthrush ( Garrulax formosus) is a Passerine Bird belonging to the Laughingthrush genus Garrulax in the Old UserPolbot. -->The Red-tailed Laughingthrush ( Garrulax milnei) is a species of Bird in the Timaliidae Auk 120(1): 1-20. The Auk is a quarterly Journal and the official publication of the American Ornithologists' Union, having been continuously published by that body since DOI: 10. A digital object identifier ( DOI) is a permanent identifier given to an Electronic document. 1642/0004-8038(2003)120[0035:MDPOBT]2. 0. CO;2 HTML fulltext without images
- Collar, N. J. & Robson, Craig (2007): Family Timaliidae (Babblers). In: del Hoyo, Josep; Elliott, Andrew & Christie, D. A. (eds. ): Handbook of Birds of the World, Volume 12 (Picathartes to Tits and Chickadees): 70-291. HBW_-_Taxonomygif|right|frame|(400 × 258 pixels file size38KBMIME typeimage/gif|HBW-Page on Taxonomy]]HBW-accounts_8-082-083_copia Lynx Edicions, Barcelona.
- Pasquet, Eric; Bourdon, Estelle; Kalyakin, Mikhail V. & Cibois, Alice (2006). The fulvettas (Alcippe), Timaliidae, Aves): a polyphyletic group. Zoologica Scripta 35, 559–566. Zoologica Scripta (ISSN 0300-3256 is a biology magazine published by Blackwell Publishing Limited, published bi-monthly doi:10.1111/j.1463-6409.2006.00253.x (HTML abstract)
- Rasmussen, Pamela C. & Anderton, J. A digital object identifier ( DOI) is a permanent identifier given to an Electronic document. Professor Pamela Cecile Rasmussen (b October 16 1959 is a prominent American ornithologist and expert on Asian Birds She was formerly a research associate C. (2005): Birds of South Asia. The Ripley Guide. Birds of South Asia redirects here For a list of the birds of South Asia see List of birds of South Asia. Smithsonian Institution and Lynx Edicions, Washington DC and Barcelona.
External links
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